Biological Oceanography

MIT-WHOI Joint Program in Oceanography/Applied Ocean Science and Engineering

Doctor of Philosophy in Biological Oceanography

Core Subject
7.470Biological Oceanography12
Foundational Statistics12
Select one of the following:
Probability and Causal Inference
Introduction to Probability and Statistics
Statistical Thinking and Data Analysis
Fundamentals of Statistics
Quantitative Methods of Data Analysis and/or Modeling12
Select one of the following: 1
An Introduction to Mathematical Ecology
Quantitative Methods and Computational Models in Neurosciences
Modeling Environmental Complexity
Modeling, Data Analysis, and Numerical Techniques for Geochemistry
Modeling the Biology and Physics of the Ocean
Graduate-Level Oceanography or Marine Science 2, 324
Elements of Modern Oceanography
Geological Oceanography
Marine Microbiology and Biogeochemistry
Marine Bioinorganic Chemistry
Marine Chemistry
Geochemistry of Marine Sediments
Marine Isotope Chemistry
Fluid Dynamics of the Atmosphere and Ocean
Introduction to Observational Physical Oceanography
Coastal Physical Oceanography
Topics in Quantitative Marine Science
Topics in Marine Ecology
Topics in Marine Physiology and Biochemistry
Topics in Biological Oceanography
Topics in Zooplankton Biology
Topics in Benthic Biology
Topics in Phytoplankton Biology
Topics in Molecular Biological Oceanography
Topics in the Behavior of Marine Animals
Topics in Marine Microbiology
Specialization36
Individually designed program leading to a specialized area of study. Options include but is not limited to benthic ecology, microbial ecology and biogeochemistry, biogeography and systematics, environmental toxicology and developmental biology, plankton ecology, marine mammals, and mathematical ecology and conservation
Thesis and Research
Pre-Thesis Research 496
Thesis (.THG subject of home department) 4288-396
Total Units480-588
1

Harvard courses OEB137 Experimental Design and Statistics for Ecology and
MCB 112 Biological Data Analysis may also be used to satisfy this requirement.

2

Other relevant courses can be requested by petition to the Joint Committee on Biological Oceanography.

3

Up to 12 of the 24 units may come from Topics subjects.

4

Biological oceanography students whose home department is Civil and Environmental Engineering register for 1.THG whether they are completing pre-thesis or thesis research, for a total of 384–492 units of 1.THG over the course of the five-year doctoral program